Selected Quarterly Financial Data (Details) (USD $)
3 Months Ended 12 Months Ended
Dec. 31, 2014
Sep. 30, 2014
Jun. 30, 2014
Mar. 31, 2014
Dec. 31, 2013
Sep. 30, 2013
Jun. 30, 2013
Mar. 31, 2013
Dec. 31, 2014
Dec. 31, 2013
Dec. 31, 2012
Statements of Operations and Comprehensive Loss                      
Net loss $ (17,962,000) $ (17,420,000) $ (12,609,000) $ (14,488,000) $ (12,531,000) $ (20,342,000) $ (19,512,000) $ (8,305,000) $ (62,479,000) $ (60,690,000) $ (69,128,000)
Net loss applicable to common stock (17,962,000) (17,420,000) (16,640,000) (19,457,000) (17,304,000) (25,090,000) (23,880,000) (11,887,000) (71,479,000) (78,161,000) (83,120,000)
Net loss per share - basic and diluted $ (0.55) $ (0.59) $ (2.22) $ (50.45) $ (44.87) $ (65.05) $ (62.59) $ (31.25)      
Weighted-average common shares outstanding - basic and diluted 32,678,459 29,746,426 7,500,148 385,664 385,688 385,688 381,525 380,352 17,699,487 383,310 368,261